The Breed History
Though one of the most popular modern dog breeds, the origins of
the Beagle are obscure. Some reports place them in England before
Roman times. Many of the early reports date to the 1300s. It is
thought that their early lineage included crosses out to other scent
hounds, and Foxhounds are variably thought to be derived from
the Beagle as a result of those crosses, or perhaps Foxhounds were
crossed back to be an early influence on beagle type. The mid 18th
century reports indicate two types of hare hunting hounds. The
modern beagle apparently derives from the North Country Beagle.
The modern American lines were first imported in the 1860s and
1880s. The breed name origins are also obscure, with deviation from
Olde English, French or Celtic being cited; the derivative may be
from roots such as "begle", "beag", or "begele".
Breeding for Function
As hunters, they are courageous and have a great deal of stamina.
They hunt equally well in packs, braced (pairs) or solo, though they
are most commonly hunted in packs. They are scent hounds and
were bred to hunt hare and rabbit. Today, they are most commonly
companion dogs.
Physical Characteristics
Height at Withers: Two distinct varieties:
1. Thirteen Inch: up to 13" high.
2. Fifteen Inch: 13-15" high (maximum height 15" in the United
States, 16" in England).
Weight: Thirteen inch variety is generally less than 20 lb (9 kg),
Fifteen inch variety is generally 20-30 lb (9-13.5 kg).
Coat: The hard, short-medium outer coat hairs may be of any
hound colors. The haircoat is dense. Black, tan and white tri-color
is very popular, also common are the red and white. Irish Spotting
is a specific spotted marking distribution. Ticking is also accepted;
blue tick and red tick being the most common of this pattern. Patch
beagle refers to a predominantly white beagle strain (with large
black, lemon or red patches generally). Dilutes (e.g., blue) in the coat
are also accepted.
Longevity: 13-15 years
Points of Conformation: The skull is broad and slightly domed, ears
are long and pendulous with round tips that hang in towards the
cheek, the leather is fine, and the ear pinna is broad. Eyes are soft
in expression, brown or hazel in color, large and wide-set. Beagles
have a moderate stop and square muzzle, minimal flews, large nose
and the nostrils start out black but fade with maturity. The neck is
medium in length and muscling with no throatiness. The back is
short, ribs are well sprung, and forelegs are straight boned. Feet are
compact and round. The tail is held high but not over the back, and
is shorter than most hound tails and is slightly curved with brush.
Recognized Behavior Issues and Traits
Reported breed characteristics include: Loyal, gentle, trustworthy,
playful, adaptable, affectionate, but can be prone to independent
thinking and will wander off if not kept on leash or fenced in.
The fence needs to be secure to prevent digging and jumping
escapes. Considered excellent with children due to a low aggression
tendency, assuming they are properly socialized. They possess a
moderate to high barking tendency. Not considered a watchdog.
Good with other pets, though may chase small ones if not
accommodated to their presence.
They have low grooming needs and low to moderate shedding
tendency. Good for city or country living. Beagles have low to
moderate exercise needs. They need close human companionship
or may howl or become destructive. Can be more of a challenge
to housebreak than some of the other breeds. Tend to gain weight
unless given adequate exercise and dietary intake is controlled.
Normal Physiologic Variations
Echocardiographic Normal Values:
Parameter Range
Weight (kg) - 5.5-12.0
LVPWD (mm) - 6-13
LVPWS (mm) - 7-17
LVD (mm) - 18-33
LVS (mm) - 8-27
FS (%) - 20-70
EF (%) - 40-100
IVSd (mm) - 5-11
IVSs (mm) - 6-12
N - 50
LVPWD, LV posterior wall dimension at end-diastole; LVPWS,
LV posterior wall thickness at end-systole; LVD, LV chamber
dimension at end-diastole; LVS, LV chamber dimension at
end-systole; FS, percent fractional shortening; EF, ejection fraction;
IVSd, interventricular septal thickness at end-diastole; IVSs,
interventricular septal thickness at end-systole; N, number of
animals.
Drug Sensitivities
None reported
Inherited Diseases
Hip Dysplasia: Polygenically inherited trait causing degenerative
joint disease and hip arthritis. OFA reports 17.8% affected.
Patella Luxation: Polygenically inherited trait causing stifle
instability and arthritis. OFA reports 1.4% affected.
Elbow Dysplasia: Polygenically inherited trait causing elbow
arthritis. Too few Beagles have been evaluated by OFA to determine
an accurate frequency.
Primary (Narrow and Open Angle) Glaucoma: Autosomal
recessive glaucoma causes bilateral increased pressure within the
eyeball between 1-2 years of age. Secondary blindness occurs due
to damage to the retina. Diagnose with tonometry and gonioscopy.
Diagnosed in 1.10% of Beagles presented to veterinary teaching
hospitals. Beagles with open angle glaucoma have increased
myocilin protein levels in the aqueous humor, with intermediate
levels in heterozygous carriers. A genetic test is not available. CERF
does not recommend breeding any Beagle with glaucoma.
Factor VII Deficiency: An autosomal recessive, mild to moderate
bleeding disorder occurs in the breed due to a mutation in the cFVII
gene. A genetic test is available.
Chondrodystrophy (Dwarfism): An autosomal recessive
dwarfism occurs in the breed. Affected dogs have a small and
disproportionate body size, a shortened neck, curved, deformed
legs, a broadened skull, and chronic arthritis. They can be diagnosed
by by x-ray stippling (i.e., punctate bright spots) of the wrist bones
under 3 months of age, or by a compressed L7 vertebrae at over 6
months. Genetic research is ongoing to identify the defective gene
at the Mark Neff lab.
Musladin-Leuke Syndrome (MLS, Chinese Beagle Syndrome):
An autosomal recessive disease that affects the development
and structure of connective tissue. It is multi-systemic, with
involvement of multiple organs, including bone, heart, skin,
and muscle. Affected dogs walk up on their toes due to tendon
contracture. They often have tighter skin with limited "scruff". They
have a flat skull, higher ear set and slanted eyes. The disease shows
variable expressivity, and affected dogs live a normal lifespan. See
the National Beagle Club and UV-Davis VGL websites for more
information. A genetic test is available.
Pyruvate Kinase Deficiency (PK): A rare, autosomal recessive
disease of red blood cells causing exercise intolerance with a
persistent, severe, and highly regenerative anemia, splenomegaly,
and progressive osteosclerosis. A genetic test is available.9,10
Tapetal Degeneration: A rare, autosomal recessive degeneration of
the tapetum occurs as a result of abnormal postnatal melanosome
development. The degeneration of the tapetum does not affect
vision and does not result in functional or structural damage to the
retina.
Disease Predispositions
Periodontal Disease: Beagles are predisposed to periodontal
disease, with 44% by two years of age, and up to 81% of older dogs
affected. The maxillary premolars are most frequently affected.
Few affected Beagles are clinically impaired, though halitosis and
gingivitis are usually present.
Distichiasis: Abnormally placed eyelashes that irritate the cornea
and conjunctiva. Can cause secondary corneal ulceration. Identified
in 17.30% of Beagles CERF examined by veterinary ophthalmologists
between 2000-2005.
Hypothyroidism: Inherited autoimmune thyroiditis. 16.5% positive
for thyroid autoantibodies based on testing at Michigan State
University.
Retinal Dysplasia: Focal folds are seen in the breed. Identified in
2.70% of Beagles CERF examined by veterinary ophthalmologists
between 2000-2005.
Cataracts: Anterior and posterior cataracts are equally represented
in the breed, and can be intermediate or punctate. Reported in
2.16% of Beagles CERF examined by veterinary ophthalmologists
between 2000-2005. CERF does not recommend breeding any
Beagle with a cataract.
Intervertebral Disc Disease (IVDD): Spinal cord disease due to
prolapsed disk material. Clinical signs include back pain, scuffing of
paws, spinal ataxia, limb weakness, and paralysis. Almost all dogs
with IVDD have calcified disk material. Beagles are significantly
over-represented with cervical IVDD. Dorn reports a 1.39x odds
ratio versus other breeds.
Cryptorchidism (Retained Testicles): Can be unilateral or bilateral.
This is a sex-limited disorder with an unknown mode of inheritance.
Reported as a breed issue on the National Beagle Club website.
Diabetes Mellitus (Sugar Diabetes): Almost exclusively female
Beagles are at increased risk of developing diabetes due to immune
mediated destruction of the pancreatic beta cells. Treat with insulin
injections, dietary management, and glucose monitoring.
Corneal Dystrophy: The breed can form oval epithelial corneal
opacities that can progressively block vision. They do not lead to
corneal edema.
Persistent Pupillary Membranes: Strands of fetal remnant
connecting; iris to iris, cornea, lens, or involving sheets of tissue.
The later three forms can impair vision, and dogs affected with
these forms should not be bred. Identified in 1.62% of Beagles CERF
examined by veterinary ophthalmologists between 2000-2005.
Prolapsed Gland of the Nictitans (Cherry Eye): This condition
occurs secondary to inflammation of the gland. Identified in 1.35%
of Beagles CERF examined by veterinary ophthalmologists between
2000-2005.
Idiopathic Epilepsy (Inherited Seizures): Control with anti-seizure
medication. Seizures generally start under one year of age. Reported
as a breed issue on the National Beagle Club website.
Pulmonic Stenosis: Narrowing of the right ventricular outflow
tract/pulmonary artery. Clinical signs in severely affected dogs
include exercise intolerance, stunting, dyspnea, syncope and ascites.
A left systolic murmur is present on auscultation. Beagles are
over-represented.
Juvenile Polyarteritis (Beagle Pain Syndrome): A naturally
occurring vasculitis and perivasculitis of unknown etiology occurs
in 6-40 month old Beagles, presenting with episodes of fever
and neck pain that last 3-7 days. Histologically, the small-to
medium-sized muscular arteries of the heart, cranial mediastinum,
and cervical spinal meninges are consistently involved.
Undetermined mode of inheritance.
Bladder and Urethral Tumors: Beagles are over-represented with
tumors of the bladder and urethra versus other breeds.
Microphthalmia with Multiple Congenital Ocular Defects:
Several congenital unilateral and bilateral microphthalmia
syndromes are recognized in the breed with associated corneal, lens,
and retinal defects. Reported in 0.54% of Beagles CERF examined
by veterinary ophthalmologists between 2000-2005. CERF does not
recommend breeding any Beagle with microphthalmia.
Deafness: Congenital deafness can be unilateral or bilateral.
Associated with extreme piebald pigmentation. Diagnose by BAER
testing. Unknown mode of inheritance.
Renal Amyloidosis: Beagles were over-represented in a study of
renal amyloidosis. Affected dogs were over 6 years of age, with a
female preponderance. Clinical signs were consistent with renal
failure. Histopathology showed moderate to severe diffuse global
glomerular amyloidosis without tubular or glomerular lesions.
Progressive Myoclonic Epilepsy (Lafora’s Disease): Older affected
Beagles show progressive, intermittent seizures that can progress
to status epilepticus. The seizures are often elicited by external
stimuli, especially a change in noise or light in the surroundings.
Electroencephalographic findings show myoclonus epilepsy.
Postmortem histopathological findings include multiple periodic
acid-Schiff-positive inclusion bodies consistent with Lafora’s
disease.
Black Hair Follicular Dysplasia, Brachygnathism, Cervical
Vertebral Instability, Cleft Lip/Palate, Copper Hepatopathy,
Cutaneous Asthenia, Demodicosis, Dilated Cardiomyopathy,
Dysfibrinogenemia, Ectodermal Defect, Elongated Soft Palate,
Factor VIII Deficiency, Globoid Cell Leukodystrophy, GM-1
Gangliosidosis, Hyperlipidemia, Hypotrichosis, IgA Deficiency,
Lissencephaly, Mitral Valve Disease, Narcolepsy, Non-Spherocytic
Hemolytic Anemia, Optic Nerve Hypoplasia, Panosteitis,
Peripheral Vestibular Disease, Prognathism, Progressive Retinal
Atrophy, Shaker Disease, Spina Bifida, Vertebral Stenosis, and
XX Sex Reversal are reported.
Isolated Case Studies
Cobalamin Malabsorption (Methylmalonic Aciduria): A
six-month-old beagle was presented with a three-month history
of failure to gain weight, lethargy, intermittent vomiting and
seizures. Laboratory results showed low serum cobalamin (Cbl)
concentrations, anaemia, leucopenia and methylmalonic aciduria.
The disorder responded to parenteral vitamin B12 therapy. This has
been identified as an autosomal recessive disorder in other breeds.
Cerebellar Cortical Abiotrophy/Ataxia: Two of five 4-1/2 month
old Beagle littermates (a male and a female) presented with
cerebellar ataxia, intension tremor, and nystagmus. Purkinje cell
degeneration in the cerebellar cortex was seen on necropsy. This has
been identified as an autosomal recessive disorder in other breeds.
Membranoproliferative Glomerulonephropathy: 5 of 7 adult
Beagles from the same litter developed polyuria, polydipsia,
proteinuria, and azotemia by 8 years of age. Membranoproliferative
glomerulonephritis was diagnosed on light and electron microscopy
of renal biopsies.
Renal Agenesis: Cases of unilateral kidney agenesis have been
documented in the literature. Compensatory renal hyperplasia
occurs in the opposite kidney, with normal kidney values.
Vetricular Septal/Conotruncal Defect: A family of Beagle dogs
was identified with congenital heart disease characterized by
ventricular septal defect and ventricular outflow abnormalities.
Breeding studies suggested an autosomal recessive mode of
inheritance.
Genetic Tests
Tests of Genotype: Direct test for MLS is available from UC-Davis
VGL.
Direct tests for PK and Factor VII deficiencies are available from
PennGen.
Direct test for Factor VII deficiency is available from VetGen.
Direct test for coat color alleles is available from VetGen.
Tests of Phenotype: CHIC certification: Required testing includes hip
radiographs, CERF eye examination a direct test for MLS, and either
a cardiac evaluation by a cardiologist, or a thyroid profile including
autoantibodies. (See CHIC website; caninehealthinfo.org).
Recommend patella evaluation and elbow radiographs.
Urine test for Cobalamine Malabsorption/Methylmalonic Aciduria is
available from PennGen.
Miscellaneous
- Breed name synonyms: English Beagle, Pocket beagle and Glove
beagle are historical nicknames for the smallest of the small variety.
- Registries: AKC, UKC, CKC, KCGB (Kennel Club of Great Britain),
ANKC (Australian National Kennel Club), NKC (National Kennel Club).
- AKC rank (year 2008): 5 (33,722 dogs registered)
- Internet resources: The National Beagle Club:
clubs.akc.org/NBC/
The Beagle Club (UK): thebeagleclub.org
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